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The Indians of the Acjachemen Nation are the original inhabitants of lands that comprise much of Orange and San Diego Counties in southern California today. They occupied the coastal region from Long Beach to Oceanside, fished off the shores of Santa Catalina and San Clemente Islands, and hunted the inland areas as far east as Lake Elsinore. Originally, they migrated into this region from the Great Basin and later experienced forced migrations within their tribal lands when Spanish, Mexican, and American invaders took their land and homes.

Wedged between two linguistically similar Indian groups, the Gabrielinos to the north and the Luiseños to the south, the Acjachemen became known as the Juaneño Indians. The Spanish discarded the Indian names of these tribes and renamed each of them after the nearby missions, including San Gabriel (Gabrielinos), San Luis Rey (Luiseños), and San Juan Capistrano (Juaneños). The Juaneños flourished before the arrival of the Europeans, living by a productive and reliable substance economy based on the hunting and gathering of naturally occurring foods and materials. The songs and stories of the tribe tell of seasonal migrations to harvest and hunt. Elaborate trade networks and kinship groups formed with other Indians in the region often dictated the migration patterns of the Juaneños. More migrations occurred within the region when the Spanish arrived in California and forced the Juaneños from their land to use it to raise crops and cattle. The Juaneños built the great mission for the invaders as diseases decimated their tribe. The Juaneños maintained their cultural identity and adapted to more changes throughout the Mexican period. The American takeover and occupation of California again forced them to sacrifice their land, relocate, and adapt to yet another new culture and language. After more than two centuries of interaction with these intruders on their land, today the large-scale migrations of the Juaneños have ended, yet their fight to retain control of ancestral lands and sacred burial grounds continues.

In terms of culture, linguistics, and origin theories, the Juaneños share similar characteristics with neighboring native groups. The Juaneño language belongs to the Takic division of the Uto-Aztecan family, and American Indians associated with the San Luis Rey and the San Juan Capistrano missions both spoke the language. These groups share cultural and linguistic traits with their Uto-Aztecan neighbors, the Cupeños, Gabrielinos, and Cahuilla Indians. Early ethnographic literature distinguishes between the Juaneños and the Luiseños, but modern academics generally agree that they were culturally one tribe. Linguistic distribution studies suggest that these people migrated into southern California from the Great Basin area in a succession of waves rather than in a single migration or drift. There is no evidence to suggest that they crossed the barrier of the Sierra Nevada to penetrate California's Central Valley during their southward migration. Their route into the region may have been via the Cajon Pass, although the San Gorgonio Pass, prominent in many of the travel songs of the Luiseño Indians, is a likely path also. Because pre–contact era ethnographical materials are scarce, no definite migration dates exist. Linguists and authors believe that these Takic-speaking groups arrived between fifteen hundred and three thousand years ago.

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